Path 1:
Speedy Nuts: throws nuts faster
Steamy Nuts: nuts pop lead and frozen bloons
Dual Tossing: throws two nuts at once
Quad Tossing: throws four nuts at once
Acorn Barrage: quickly shoots ten nuts at once
Path 2:
Hard Nuts: nuts have two pierce
Pointed Nuts: nuts pop two layers at once
Nut Impact: nuts pop four layers at once
Berserk: ability instantly sends squirrel into hypersonic rage
Meltdown: squirrel now throws five nuts in rage
Path 3:
Nut Launch: nuts bounce from one bloon to another
Building Anger: squirrel goes angry every 100 bloons popped
Bloon Rage: now gets angry every 50 bloons popped
True Fury: gets angry every 20 bloons popped
Temper Tantrum: has 10 pierce whenever he gets angry